Cochemiea

Cochemiea is a genus of cactus.[2] It has previously been synonymized with Mammillaria, but molecular phylogenetic studies have shown that when broadly circumscribed, Mammillaria is not monophyletic, and Cochemiea has been accepted as a separate genus.[3]

Cochemiea
Cochemiea macdougallii
Scientific classification
Kingdom: Plantae
Clade: Tracheophytes
Clade: Angiosperms
Clade: Eudicots
Order: Caryophyllales
Family: Cactaceae
Subfamily: Cactoideae
Tribe: Cacteae
Genus: Cochemiea
(K.Brandegee) Walton[1]
Species

See text.

Taxonomy

A molecular phylogenetic study of the "mammilloid clade" showed that it consisted of four monophyletic groups, which the authors re-circumscribed into four genera: Cumarinia; Mammillaria, with a reduced number of species; Coryphantha, expanded to include species previously placed in Mammillaria and Escobaria; and Cochemiea, expanded to include a large number of species previously placed in Mammillaria.[3]

Mammilloid clade
 

Cumarinia

 
 
 

Mammillaria s.s.

 
 
 

Coryphantha s.l.

 
 

Cochemiea s.l.

 
 
 
 

Species

In 2021, Breslin, Wojciechowski and Majure placed the following species in the genus, some already placed there and some moved from Mammillaria, Neomammillaria and Ortegocactus.[3] As of March 2022, Plants of the World Online did not include in the genus the species moved by Breslin et al.[2]

  • Cochemiea albicans (Britton & Rose) P.B.Breslin & Majure, syn. Mammillaria albicans (Britton & Rose) A.Berger
  • Cochemiea angelensis (R.T.Craig) P.B.Breslin & Majure, syn. Mammillaria angelensis R.T.Craig
  • Cochemiea armillata (K.Brandegee) P.B.Breslin & Majure, syn. Mammillaria armillata K.Brandegee
  • Cochemiea barbata (Engelm.) Doweld
  • Cochemiea blossfeldiana (Boed.) P.B.Breslin & Majure, syn. Mammillaria blossfeldiana Boed.
  • Cochemiea boolii (G.E.Linds.) P.B.Breslin & Majure,, syn. Mammillaria boolii G.E.Linds.
  • Cochemiea bullardiana (H.E.Gates) P.B.Breslin & Majure,, syn. Neomammillaria bullardiana H.E.Gates
  • Cochemiea capensis (H.E.Gates) Doweld
  • Cochemiea cerralboa (Britton & Rose) P.B.Breslin & Majure, syn. Neomammillaria cerralboa Britton & Rose
  • Cochemiea conoidea (DC.) P.B.Breslin & Majure, syn. Mammillaria conoidea DC.
  • Cochemiea dioica (K.Brandegee) Doweld
  • Cochemiea estebanensis (G.E.Linds.) P.B.Breslin & Majure,, syn. Mammillaria estebanensis G.E.Linds.
  • Cochemiea fraileana (Britton & Rose) P.B.Breslin & Majure, syn. Neomammillaria fraileana Britton & Rose, Mammillaria albicans subsp. fraileana (Britton & Rose) D.R.Hunt
  • Cochemiea goodridgei (Scheer ex Salm-Dyck) P.B.Breslin & Majure, syn. Mammillaria goodridgei Scheer ex Salm-Dyck
  • Cochemiea grahamii (Engelm.) Doweld
  • Cochemiea guelzowiana (Werderm.) P.B.Breslin & Majure, syn. Mammillaria guelzowiana Werderm.
  • Cochemiea halei (K.Brandegee) Walton
  • Cochemiea hutchisoniana (H.E.Gates) P.B.Breslin & Majure, syn. Neomammillaria hutchisoniana H.E.Gates, Mammillaria hutchisoniana (H.E.Gates) Boed.
  • Cochemiea insularis (H.E.Gates) P.B.Breslin & Majure, syn. Mammillaria insularis H.E.Gates ex Shurly
  • Cochemiea macdougallii (Alexander) P.B.Breslin & Majure, syn. Ortegocactus macdougallii Alexander
  • Cochemiea mainiae (K.Brandegee) P.B.Breslin & Majure, syn. Mammillaria mainiae K.Brandegee
  • Cochemiea maritima H.E.Gates ex. Shurly
  • Cochemiea multidigitata (Radley & G.E.Linds.) P.B.Breslin, syn. Mammillaria multidigitata Radley & G.E.Linds
  • Cochemiea palmeri (J.M.Coult.) P.B.Breslin & Majure, syn. Mammillaria neopalmeri R.T.Craig
  • Cochemiea phitauiana (E.M.Baxter) Doweld
  • Cochemiea pondii Walton
  • Cochemiea poselgeri (Hildm.) Britton & Rose
  • Cochemiea saboae (Glass) Doweld
  • Cochemiea schumannii (Hildm.) P.B.Breslin & Majure, syn. Mammillaria schumannii Hildm.
  • Cochemiea setispina Walton
  • Cochemiea sheldonii (Britton & Rose) Doweld
  • Cochemiea swinglei (Britton & Rose) Doweld
  • Cochemiea tetrancistra (Engelm.) P.B.Breslin & Majure, syn. Mammillaria tetrancistra Engelm.
  • Cochemiea theresae (Cutak) Doweld
  • Cochemiea thomasii García-Mor. & al.
  • Cochemiea thornberi (Orcutt) P.B.Breslin & Majure, syn. Mammillaria thornberi Orcutt
  • Cochemiea viridiflora (Britton & Rose) P.B.Breslin & Majure, syn. Neomammillaria viridiflora Britton & Rose
  • Cochemiea wrightii (Engelm.) Doweld
  • Cochemiea angelensis
  • Cochemeia boolii
  • Cochemeia mainiae
  • Cochemiea tetrancistra

References
  1. "Cochemiea (K.Brandegee) Walton", The International Plant Names Index, retrieved 2022-03-05
  2. "Cochemiea (K.Brandegee) Walton", Plants of the World Online, Royal Botanic Gardens, Kew, retrieved 2022-03-05
  3. Breslin, Peter B.; Wojciechowski, Martin F. & Majure, Lucas C. (2021), "Molecular phylogeny of the Mammilloid clade (Cactaceae) resolves the monophyly of Mammillaria", Taxon, 70 (2): 308–323, doi:10.1002/tax.12451, S2CID 233885318
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