2022 in paleomalacology

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This list of fossil molluscs described in 2022 is a list of new taxa of fossil molluscs that were described during the year 2022, as well as other significant discoveries and events related to molluscan paleontology that occurred in 2022.

Ammonites

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Acanthohoplites albedocostatus[1]	Sp. nov	In press	Matsukawa	Early Cretaceous		Japan
Acantholytoceras costaevariavilis[1]	Sp. nov	In press	Matsukawa	Early Cretaceous		Japan
Dactylioceras kopiki[2]	Sp. nov	Valid	Rogov in Rogov & Lutikov	Early Jurassic (Toarcian)		Canada ( Northwest Territories) Norway United States ( Alaska) Russia?	A member of the family Dactylioceratidae.
Epitornoceras transmediterraneum[3]	Sp. nov	Valid	Korn & Bockwinkel	Devonian (Givetian)		Morocco	A member of the family Tornoceratidae.
Kutatissites sudai[1]	Sp. nov	In press	Matsukawa	Early Cretaceous		Japan
Lentitornoceras[3]	Gen. et sp. nov	Valid	Korn & Bockwinkel	Devonian (Givetian)	Red Ironstone Formation	Germany	A member of the family Tornoceratidae. The type species is L. materni.
Megatyloceras leteilensis[4]	Sp. nov	Valid	Delanoy et al.	Early Cretaceous (Aptian)		France	A member of the family Douvilleiceratidae.
Nejdia arabica[5]	Sp. nov	Valid	Gelin	Early Jurassic (Toarcian)	Marrat Formation	Saudi Arabia
Paracrochordiceras watanabei[6]	Sp. nov	Valid	Ehiro	Middle Triassic (Anisian)	Fukkoshi Formation	Japan	A member of the family Acrochordiceratidae.
Paradanubites ozashiense[6]	Sp. nov	Valid	Ehiro	Middle Triassic (Anisian)	Fukkoshi Formation	Japan	A member of Danubitoidea belonging to the family Danubitidae.
Parahedenstroemia? tatjanae[7]	Sp. nov	Valid	Đaković, Krystyn & Sudar	Early Triassic		Montenegro Russia?	A member of the family Hedenstroemiidae.
Parapopanoceras involutum[6]	Sp. nov	Valid	Ehiro	Middle Triassic (Anisian)	Fukkoshi Formation	Japan	A member of Ceratitida belonging to the group Megaphyllitoidea and the family Parapopanoceratidae.
Paucitornoceras[3]	Gen. et comb. nov	Valid	Korn & Bockwinkel	Devonian	Red Ironstone Formation	Germany United States ( New York)	A member of the family Tornoceratidae. The type species is "Goniatites" paucistriatus d`Archiac & de Verneuil (1842); genus also includes "Tornoceras" rhysum Clarke (1898).
Procheloniceras yajimai[1]	Sp. nov	In press	Matsukawa	Early Cretaceous		Japan
Pseudosaynella pacifica[1]	Sp. nov	In press	Matsukawa	Early Cretaceous		Japan
Psilokhvalynites[6]	Gen. et sp. nov	Valid	Ehiro	Middle Triassic (Anisian)	Fukkoshi Formation	Japan	A member of Dinaritoidea belonging to the family Khvalynitidae. The type species is P. takaizumii.
Shasticrioceras nakazawai[1]	Sp. nov	In press	Matsukawa	Early Cretaceous		Japan

Ammonite research

A study on the morphological change through ontogeny of beyrichitine ammonites from the Anisian of Nevada is published by Bischof, Schlüter & Lehmann (2022).[8]
A study aiming to determine whether the environmental changes around the Triassic–Jurassic boundary influenced development of septal thickness in ammonites, as well aiming to determine the relation between septal thickness and paleolatitudinal occurrences, and whether ontogenetic septal thickness-trajectories reflected paleogeographic origin or phylogenetic relationships of ammonites, is published by Weber et al. (2022).[9]
A study on the hydrodynamice properties of shells of Late Triassic and Early Jurassic ammonites is published by Hebdon et al. (2022).[10]
A study on changes of septal spacing between succeeding chambers of the shell of Gaudryceras tenuiliratum during its ontogeny is published by Kawakami, Uchiyama & Wani (2022).[11]

Other cephalopods

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Arctoteuthis angusta[12]	Sp. nov	Valid	Efremenko	Early Cretaceous		Russia	A belemnite belonging to the family Cylindroteuthididae.
Boreioteuthis mirifica[12]	Sp. nov	Valid	Efremenko	Early Cretaceous		Russia	A belemnite belonging to the family Cylindroteuthididae.
Domatoceras tuckeri[13]	Sp. nov	Valid	Niko, Mapes & Seuss	Carboniferous (late Pennsylvanian)	Graham Formation	United States ( Texas)	A member of the family Grypoceratidae.
Endolobus sturgeoni[13]	Sp. nov	Valid	Niko, Mapes & Seuss	Carboniferous (late Pennsylvanian)	Graham Formation	United States ( Texas)	A member of the family Koninckioceratidae.
Holmiceras havelense[14]	Sp. nov	Valid	Aubrechtová & Korn	Ordovician (Darriwilian)	Upper Red Orthoceratite Limestone	Germany
Lispoceras orbis[15]	Sp. nov	Valid	Korn, Miao & Bockwinkel	Early Carboniferous		Algeria	A member of the family Trigonoceratidae.
Lituites baculus[14]	Sp. nov	Valid	Aubrechtová & Korn	Middle Ordovician		Sweden
Lituites bottkei[14]	Sp. nov	Valid	Aubrechtová & Korn	Middle Ordovician		Poland Sweden
Lituites clavis[14]	Sp. nov	Valid	Aubrechtová & Korn	Ordovician (Darriwilian)	Upper Grey Orthoceratite Limestone	Germany
Lituites dewitzi[14]	Sp. nov	Valid	Aubrechtová & Korn	Ordovician (Darriwilian)	Upper Grey Orthoceratite Limestone	Germany
Lituites kruegeri[14]	Sp. nov	Valid	Aubrechtová & Korn	Ordovician (Darriwilian)	Upper Grey Orthoceratite Limestone	Germany
Lituites nebeni[14]	Sp. nov	Valid	Aubrechtová & Korn	Ordovician (Darriwilian)	Upper Grey Orthoceratite Limestone	Germany
Maccoyoceras pentagonum[15]	Sp. nov	Valid	Korn, Miao & Bockwinkel	Early Carboniferous		Algeria	A member of the family Trigonoceratidae.
Metacoceras quadratum[13]	Sp. nov	Valid	Niko, Mapes & Seuss	Carboniferous (late Pennsylvanian)	Graham Formation	United States ( Texas)	A member of the family Tainoceratidae.
Planetoceras destrictum[15]	Sp. nov	Valid	Korn, Miao & Bockwinkel	Early Carboniferous		Algeria	A member of the family Trigonoceratidae.
Planetoceras transforme[15]	Sp. nov	Valid	Korn, Miao & Bockwinkel	Early Carboniferous		Algeria	A member of the family Trigonoceratidae.
Pleuronautilus chisatoi[16]	Sp. nov	Valid	Ehiro	Permian (Wordian)	Takakurayama Formation	Japan	A member of the family Tainoceratidae.
Rhynchorthoceras kranepuhlense[14]	Sp. nov	Valid	Aubrechtová & Korn	Ordovician (Darriwilian)	Upper Red Orthoceratite Limestone	Germany Poland
Rhynchorthoceras rugium[14]	Sp. nov	Valid	Aubrechtová & Korn	Ordovician (Darriwilian)	Upper Red Orthoceratite Limestone	Germany
Rineceras multituberculatum[15]	Sp. nov	Valid	Korn, Miao & Bockwinkel	Early Carboniferous		Algeria	A member of the family Trigonoceratidae.
Rineceras rectangulatum[15]	Sp. nov	Valid	Korn, Miao & Bockwinkel	Early Carboniferous		Algeria	A member of the family Trigonoceratidae.
Solenochilus jackense[13]	Sp. nov	Valid	Niko, Mapes & Seuss	Carboniferous (late Pennsylvanian)	Graham Formation	United States ( Texas)	A member of the family Solenochilidae.
Syllipsimopodi[17]	Gen. et sp. nov	Valid	Whalen & Landman	Carboniferous (Serpukhovian)	Bear Gulch Limestone	United States ( Montana)	A member of Coleoidea belonging to the clade Vampyropoda. The type species is S. bideni, named for Joe Biden.[18]
Thrincoceras devolvere[15]	Sp. nov	Valid	Korn, Miao & Bockwinkel	Early Carboniferous		Algeria	A member of the family Trigonoceratidae.
Tohokubelus[19]	Gen. et sp. nov	Valid	Niko & Ehiro	Early Triassic (Olenekian)	Osawa Formation	Japan	A sinobelemnitid belemnite. Genus includes new species T. takaizumii.
Trilacinoceras filix[14]	Sp. nov	Valid	Aubrechtová & Korn	Ordovician (Darriwilian)	Upper Grey Orthoceratite Limestone	Germany	A member of the family Lituitidae.
Trilacinoceras knoefleri[14]	Sp. nov	Valid	Aubrechtová & Korn	Ordovician (Darriwilian)	Upper Grey Orthoceratite Limestone	Germany	A member of the family Lituitidae.
Vestinautilus concinnus[15]	Sp. nov	Valid	Korn, Miao & Bockwinkel	Early Carboniferous		Algeria	A member of the family Trigonoceratidae.
Vestinautilus padus[15]	Sp. nov	Valid	Korn, Miao & Bockwinkel	Early Carboniferous		Algeria	A member of the family Trigonoceratidae.

Other cephalopod research

A study on the phylogenetic relationships of Cambrian and Ordovician cephalopods is published by Pohle et al. (2022).[20]
A study on the identity of enigmatic black amorphous fossils often associated with belemnoid remains, based on the analysis of fossil material from the Upper Triassic (Carnian) Polzberg Lagerstätte (Austria), is published by Lukeneder & Lukeneder (2022), who interpret the enigmatic fossils as likely representing the preservation of a mineralized and secondarily carbonized cephalic-ocular-arm-cartilage complex of the belemnoid Phragmoteuthis bisinuata.[21]
Fossil material of "Kummelonautilus" taiwanum is described from the Miocene Houdongkeng Formation (Taiwan) by Goedert, Kiel & Tsai (2022), who provisionally transfer this species to the genus Nautilus, and review the fossil record of the genus Nautilus in the Indo-Pacific region.[22]

Bivalves

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Crassatella (Bathytormus) ignava[23]	Sp. nov	Valid	Berezovsky	Eocene	Malinovskaya Formation	Ukraine	A member of the family Crassatellidae.
Crassatella lassa[24]	Sp. nov	Valid	Berezovsky	Late Eocene		Ukraine
Myonia komiensiformis[25]	Sp. nov	Valid	Biakov	Permian		Russia	A member of Pholadomyida belonging to the family Sanguinolitidae.
Myonia kutygini[25]	Sp. nov	Valid	Biakov	Permian		Russia	A member of Pholadomyida belonging to the family Sanguinolitidae.
Palaeomutela (Palaeanodonta) berrutii[26]	Sp. nov	Valid	Silantiev et al.	Permian (Kungurian)	Collio Formation	Italy	A member of Actinodontida belonging to the family Palaeomutelidae.
Palaeomutela (Palaeanodonta) guncinaensis[26]	Sp. nov	Valid	Silantiev et al.	Permian (Kungurian)	Guncina Formation	Italy	A member of Actinodontida belonging to the family Palaeomutelidae.

Bivalve research

A study on the impact of the Triassic–Jurassic extinction event on the body-size distribution of bivalves, based on data from three study sites in the United Kingdom, is published by Opazo & Twitchett (2022).[27]

Gastropods

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Ampezzopleura barremica[28]	Sp. nov	Valid	Isaji, Haga & Kashiwagi	Early Cretaceous (Barremian)	Kimigahama Formation	Japan
Anisocycla subcylindrica[29]	Sp. nov	Valid	Pavia et al.	Pliocene		Italy	A member of the family Murchisonellidae.
Antiphora aurora[28]	Sp. nov	Valid	Isaji, Haga & Kashiwagi	Early Cretaceous (Barremian)	Kimigahama Formation	Japan
Baylea tenera[30]	Sp. nov	Valid	Karapunar et al.	Carboniferous (Pennsylvanian)	Boggy Formation	United States ( Oklahoma)	A member of the family Eotomariidae.
Choshipleura[28]	Gen. et sp. nov	Valid	Isaji, Haga & Kashiwagi	Early Cretaceous (Barremian)	Kimigahama Formation	Japan	Genus includes new species C. striata.
"Clavatula" ariejansseni[31]	Sp. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Serravallian)	Týrtar Formation	Turkey	A member of the family Clavatulidae.
"Clavatula" atatuerki[31]	Sp. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Serravallian)	Týrtar Formation	Turkey	A member of the family Clavatulidae.
"Clavatula" irisae[31]	Sp. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)	Baden Formation	Austria Slovenia	A member of the family Clavatulidae.
"Clavatula" jarzynkae[31]	Nom. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)	Grund Formation	Austria Poland	A member of the family Clavatulidae; a replacement name for Clavatula auingeri Finlay (1927).
Clavatula sorini[31]	Sp. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)	Dej Formation	Romania Italy?	A species of Clavatula.
Columbellaria rara[32]	Sp. nov	Valid	Gründel, Hostettler & Menkveld-Gfeller	Late Jurassic (Oxfordian)	St-Ursanne Formation	Switzerland	A member of the family Colombellinidae.
Conus damianakisi[33]	Sp. nov	Valid	Psarras, Merle & Koskeridou	Miocene (Tortonian)		Greece	A species of Conus.
Conus davolii[33]	Sp. nov	Valid	Psarras, Merle & Koskeridou	Miocene (Tortonian)		Greece Italy Hungary?	A species of Conus.
Conus dellabellai[29]	Sp. nov	Valid	Pavia et al.	Pliocene		Italy	A species of Conus.
Conus ictini[33]	Sp. nov	Valid	Psarras, Merle & Koskeridou	Miocene (Serravallian and Tortonian)		Greece Turkey	A species of Conus.
Conus lauriatragei[33]	Sp. nov	Valid	Psarras, Merle & Koskeridou	Miocene (Langhian to Tortonian)		Greece Hungary	A species of Conus.
Conus moissettei[33]	Sp. nov	Valid	Psarras, Merle & Koskeridou	Miocene (Tortonian)		Greece	A species of Conus.
Conus villalvernensis[29]	Sp. nov	Valid	Pavia et al.	Pliocene		Italy	A species of Conus.
Crepidula bellardii[29]	Sp. nov	Valid	Pavia et al.	Pliocene		Italy	A species of Crepidula.
Cryptoptyxis? fortiter[32]	Sp. nov	Valid	Gründel, Hostettler & Menkveld-Gfeller	Late Jurassic (Oxfordian)	St-Ursanne Formation	Switzerland	A member of the family Maoraxidae.
Dictyotomaria turrisbabel[30]	Sp. nov	Valid	Karapunar et al.	Carboniferous (Pennsylvanian)	Caddo Creek Formation	United States ( Texas)	A member of the family Phymatopleuridae.
Diempterus? multicostatus[32]	Sp. nov	Valid	Gründel, Hostettler & Menkveld-Gfeller	Late Jurassic (Oxfordian)	St-Ursanne Formation	Switzerland	A member of the family Aporrhaidae.
Eirlysella[30]	Gen. et sp. et comb. nov	Valid	Karapunar et al.	Carboniferous (Pennsylvanian)	Boggy Formation	United States ( Oklahoma)	A member of the family Luciellidae. The type species is E. buckhornensis; genus also includes "Trochus" hissingerianus de Koninck (1843), "Pleurotomaria" squamula Phillips (1836) and "Luciellina" ocultabanda Kues & Batten (2001).
Exelissa ursannella[32]	Sp. nov	Valid	Gründel, Hostettler & Menkveld-Gfeller	Late Jurassic (Oxfordian)	St-Ursanne Formation	Switzerland	A member of Cerithioidea belonging to the family Cryptaulacidae.
Granulatocincta[31]	Gen. et comb. et 3 sp. nov	Valid	Harzhauser, Landau & Janssen	Miocene and Pliocene		Austria Bulgaria Czech Republic France Hungary Italy Romania Slovakia Spain Turkey	A member of the family Clavatulidae. The type species is "Pleurotoma" granulato-cincta Münster in Goldfuss (1841); genus also includes new species G. callim, G. theoderichi and G. pelliscrocodili, as well as "Pleurotoma (Clavatula)" angelae Hoernes & Auinger (1891), "Clavatula" contorta Švagrovský (1958), "Pleurotoma (Clavatula)" nataliae Hoernes & Auinger (1891), "Pleurotoma" schreibersi Hörnes (1854), "Clavatula" calcarai Bellardi (1877), "Clavatula" curionii Bellardi (1877), "Clavatula" pellegrinii Simonelli (1896), "Pleurotoma" sotterii Michelotti (1847), "Clavatula" turbinata Bellardi (1877), "Clavatula" turgidula Bellardi (1877), "Clavatula" vigolensis Bellardi (1877), "Clavatula" rugata Bellardi (1877), "Murex" rusticus Brocchi (1814), "Pleurotoma" capgrandi Tournouër (1873), "Pleurotoma" obtruta Millet (1865), "Clavatula" delgadoi Vera-Peláez & Lozano-Francisco (2001) and possibly "Clavatula" boreointerrupta Kautsky (1925).
Loriolotrema? nodosa[32]	Sp. nov	Valid	Gründel, Hostettler & Menkveld-Gfeller	Late Jurassic (Oxfordian)	St-Ursanne Formation	Switzerland	A member of the family Brachytrematidae.
Lyria estotiensis[34]	Sp. nov	Valid	Bail, Ledon & Pacaud	Oligocene (Chattian)		France	A species of Lyria.
Lyria lozoueti[34]	Sp. nov	Valid	Bail, Ledon & Pacaud	Miocene (Aquitanian)		France	A species of Lyria.
Megaclavatula[31]	Gen. et 2 sp. et comb. nov	Valid	Harzhauser, Landau & Janssen	Miocene		Austria Czech Republic Hungary Poland Romania Slovakia Slovenia Ukraine Italy?	A member of the family Clavatulidae. The type species is M. grunerti; genus also includes new species M. pilleri, as well as "Pleurotoma (Clavatula)" amaliae Hoernes & Auinger (1891), "Pleurotoma (Clavatula)" antoniae Hoernes & Auinger (1891), "Pleurotoma (Clavatula)" evae Hoernes & Auinger (1891), "Pleurotoma" laevigatum Eichwald (1830), "Perrona" nemethi Kovács & Vicián (2021), "Pleurotoma" neudorfensis Schaffer (1898), "Pleurotoma tuberculosa" polonica Pusch (1837), "Pleurotoma (Clavatula) asperulata" var. subsculpta Schaffer (1912), "Pleurotoma" ernesti Toula (1901), "Pleurotoma" asperulata Lamarck (1822), Clavatula saubrigiana laurensii Peyrot (1931) and possibly "Pleurotoma (Clavatula)" szontaghi Strausz (1926), "Clavatula" taurofusulata Sacco (1890), "Clavatula" subdepressa Sacco (1890) and "Clavatula" tortonica Peyrot (1931).
Metacerithium boshuae[28]	Sp. nov	Valid	Isaji, Haga & Kashiwagi	Early Cretaceous (Barremian)	Kimigahama Formation	Japan	A member of the family Metacerithiidae.
Neoperrona[31]	Gen. et comb. et sp. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)		Croatia Hungary Italy	A member of the family Clavatulidae. The type species is "Perrona" harzhauseri Kovács & Vicián (2021); genus also includes new species N. zoltanorum, as well as "Clavatula" taurinensis Bellardi (1877).
Neuburgensia[32]	Gen. et comb. nov	Valid	Gründel, Hostettler & Menkveld-Gfeller	Late Jurassic (Oxfordian to Tithonian)		Germany Switzerland	Possibly a member of Cerithioidea. The type species is "Gymnocerithium" concavum Janicke (1966); genus also includes "Cerithium" amabile Zittel (1873), "Gymnocerithium"? convexoconcavum Gründel, Keupp & Lang (2019), "Cerithium" crenato-cinctum Zittel (1873), "Cerithium" involvens Zittel (1873) and "Cerithium" perrotundum Cossmann (1913).
Odetta chirlii[29]	Sp. nov	Valid	Pavia et al.	Pliocene		Italy	A species of Odetta.
Olegia[31]	Gen. et sp. et comb. nov	Junior homonym	Harzhauser, Landau & Janssen	Miocene (Burdigalian to Tortonian)		Austria Bosnia and Herzegovina Bulgaria Croatia Czech Republic Italy Moldova Poland Romania Slovakia Ukraine	A member of the family Clavatulidae. The type species is O. mandici; genus also includes "Pleurotoma (Clavatula)" agathae Hoernes & Auinger (1891), "Pleurotoma (Clavatula)" brigittae Hoernes & Auinger (1891), "Pleurotoma" doderleini Hörnes (1854), "Pleurotoma (Clavatula)" dorotheae Hoernes & Auinger (1891), "Pleurotoma" rumana Simionescu & Barbu (1940), Clavatula schreibersi szokolyensis Strausz (1960), "Pleurotoma" winterlingensis Quenstedt (1884), "Clavatula" aradasi Bellardi (1877), "Clavatula" baccifera Bellardi (1877), "Clavatula" consularis Bellardi (1877) and "Clavatula" stazzanensis Bellardi (1877). The generic name is preoccupied by Olegia Shaposhnikov (1979).
Oligoptyxis khorasanica[35]	Sp. nov	Valid	Guzhov & Nazemi in Raisossadat et al.	Early Cretaceous (Aptian)		Iran	A member of the family Nerineidae.
Ondina curta[29]	Sp. nov	Valid	Pavia et al.	Pliocene		Italy	A species of Ondina.
Ondina elongata[29]	Sp. nov	Valid	Pavia et al.	Pliocene		Italy	A species of Ondina.
Ondina pseudovitrea[29]	Sp. nov	Valid	Pavia et al.	Pliocene		Italy	A species of Ondina.
Ovalotrema[32]	Gen. et comb. nov	Valid	Gründel, Hostettler & Menkveld-Gfeller	Late Jurassic (Oxfordian)		France Switzerland	A member of the family Brachytrematidae. The type species is "Turbo" cotteausius d’Orbigny (1853); genus might also include "Turbo" greppini De Loriol in De Loriol & Koby (1895).
Paragoniozona ornata[30]	Nom. nov	Valid	Karapunar et al.	Carboniferous (Pennsylvanian)	Lazy Bend Formation	United States ( Texas)	A member of the family Phymatopleuridae; a replacement name for Pleurotomaria aspera Girty (1934).
Paragoniozona yanceyi[30]	Sp. nov	Valid	Karapunar et al.	Carboniferous (Pennsylvanian)	Boggy Formation	United States ( Oklahoma)	A member of the family Phymatopleuridae.
Perrona grossi[31]	Nom. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)	Florian Formation	Austria	A species of Perrona; a replacement name for Pleurotoma (Clavatula) auingeri Hilber (1879).
Perrona ilonae[31]	Nom. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)	Sámsonháza Formation	Hungary Ukraine	A species of Perrona; a replacement name for Clavatula vindobonensis nodosa Csepreghy-Meznerics (1954).
Perrona koeberli[31]	Sp. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)	Grund Formation	Austria	A species of Perrona.
Perrona loetschi[31]	Sp. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)	Grund Formation	Austria	A species of Perrona.
Perrona wanzenboecki[31]	Nom. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)	Baden Formation	Austria	A species of Perrona; a replacement name for Pleurotoma concinna Handmann (1883).
Peruvispira kanchanaburiensis[36]	Sp. nov	In press	Karapunar, Nützel & Ketwetsuriya	Permian	Ratburi Group	Thailand	A member of the family Goniasmatidae.
Peruvispira oklahomaensis[30]	Sp. nov	Valid	Karapunar et al.	Carboniferous (Pennsylvanian)	Gene Autry Formation	United States ( Oklahoma)	A member of the family Goniasmatidae.
Phymatopleura conica[30]	Sp. nov	Valid	Karapunar et al.	Carboniferous (Pennsylvanian)	Gene Autry Formation	United States ( Oklahoma)	A member of the family Phymatopleuridae.
Phymatopleura girtyi[30]	Sp. nov	Valid	Karapunar et al.	Carboniferous (Pennsylvanian)	Wewoka Formation	United States ( Oklahoma)	A member of the family Phymatopleuridae.
Pseudomelania yamadai[28]	Sp. nov	Valid	Isaji, Haga & Kashiwagi	Early Cretaceous (Barremian)	Kimigahama Formation	Japan
Purpuripullina[35]	Gen. et comb. nov	Valid	Guzhov & Raisossadat in Raisossadat et al.	Cretaceous		Iran Lebanon	The type species is "Natica" scalaris Conrad (1852).
Pusionella hofmanni[31]	Sp. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)	Dej Formation	Romania	A species of Pusionella.
Saalensia lorioli[32]	Sp. nov	Valid	Gründel, Hostettler & Menkveld-Gfeller	Late Jurassic (Oxfordian)	St-Ursanne Formation	Switzerland	A member of the family Brachytrematidae.
Shansiella (Oklahomaella) globilineata[30]	Sp. nov	Valid	Karapunar et al.	Carboniferous (Pennsylvanian)	Gene Autry Formation	United States ( Oklahoma)	A member of the family Portlockiellidae.
Spinatus ruizi[37]	Sp. nov	Valid	Liverani & Wieneke	Miocene (Langhian)		Indonesia	A member of the family Strombidae.
Spiroscala quasipulchra[30]	Nom. nov	Valid	Karapunar et al.	Permian		United States	A member of the family Eotomariidae; a replacement name for Euconospira pulchra Batten (1958).
Spiroscala shwedagoniformis[30]	Sp. nov	Valid	Karapunar et al.	Carboniferous (Pennsylvanian)	Gene Autry Formation	United States ( Oklahoma)	A member of the family Eotomariidae.
Striopusionella[31]	Gen. et comb. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)		Hungary	A member of the family Clavatulidae. The type species is "Clavatula" szekelyhidiae Kovács & Vicián (2021).
Stuoraxis kasei[28]	Sp. nov	Valid	Isaji, Haga & Kashiwagi	Early Cretaceous (Barremian)	Kimigahama Formation	Japan	A member of the family Stuoraxidae.
Tomellana aueri[31]	Sp. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)	Baden Formation	Austria Bulgaria Hungary Romania Slovakia Slovenia	A species of Tomellana.
Tomellana dulaii[31]	Sp. nov	Valid	Harzhauser, Landau & Janssen	Miocene (Langhian)	Dej Formation	Hungary Romania	A species of Tomellana.
Worthenia (Yochelsonospira) kuesi[30]	Sp. nov	Valid	Karapunar et al.	Carboniferous (Pennsylvanian)	Wetumka Formation	United States ( Oklahoma)	A species of Worthenia.

Gastropod research

A study aiming to determine the drivers of diversification for European freshwater gastropods over the past 100 million years is published by Neubauer et al. (2022).[38]
A study aiming to determine the factors which caused the diversification of European freshwater gastropods in the Late Cretaceous is published by Neubauer & Harzhauser (2022).[39]
Revision of the fossil material of Ancistrolepis from the Eocene (Ypresian) Llajas Formation (California, United States), representing the oldest occurrence of this genus reported to date, is published by Squires (2022).[40]

Other molluscs

New taxa

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Calloplax roederi[41]	Sp. nov	Valid	Vendrasco, Powell & LaFollette	Miocene		United States ( California)	A chiton.
Chiton solaris[41]	Sp. nov	Valid	Vendrasco, Powell & LaFollette	Miocene		United States ( California)	A chiton.
Dorispira avannga[42]	Sp. nov	Valid	Peel & Kouchinsky	Cambrian (Wuliuan)	Henson Gletscher Formation	Greenland	A helcionellid.
Dorispira septentrionalis[42]	Sp. nov	Valid	Peel & Kouchinsky	Cambrian (Wuliuan)	Henson Gletscher Formation	Greenland	A helcionellid.
Dorispira tavsenensis[42]	Sp. nov	Valid	Peel & Kouchinsky	Cambrian (Wuliuan)	Henson Gletscher Formation	Greenland	A helcionellid.
Dorispira tippik[42]	Sp. nov	Valid	Peel & Kouchinsky	Cambrian (Wuliuan)	Henson Gletscher Formation	Greenland	A helcionellid.
Erugoconus[42]	Gen. et sp. nov	Valid	Peel & Kouchinsky	Cambrian (Wuliuan)	Henson Gletscher Formation	Greenland	A member of Helcionelloida of uncertain affinities. The type species is E. acuminatus.
Ischnochiton loureiroi[43]	Sp. nov	Valid	Dell'Angelo et al.	Pliocene	Mondego Basin	Portugal	A chiton, a species of Ischnochiton.
Lepidochitona rochae[43]	Sp. nov	Valid	Dell'Angelo et al.	Pliocene	Mondego Basin	Portugal	A chiton, a species of Lepidochitona.
Scenella? siku[42]	Sp. nov	Valid	Peel & Kouchinsky	Cambrian (Wuliuan)	Henson Gletscher Formation	Greenland
Sermeqiconus[42]	Gen. et comb. nov	Valid	Peel & Kouchinsky	Cambrian	Aftenstjernesø Formation	Greenland	A helcionellid. The type species is "Figurina" polaris Peel (2021).
Tavseniconus[42]	Gen. et sp. nov	Valid	Peel & Kouchinsky	Cambrian (Wuliuan)	Henson Gletscher Formation	Greenland	A member of Helcionelloida of uncertain affinities. The type species is T. erectus.
Vendrascospira[42]	Gen. et 2 sp. nov	Valid	Peel & Kouchinsky	Cambrian (Wuliuan)	Henson Gletscher Formation	Greenland	A helcionellid. The type species is V. troelseni; genus also includes V. frykmani.

Other mollusc research

Revision of the oldest known specimens of Jinonicella kolebabai from the Middle Ordovician strata of Ukraine and Belarus is published by Gubanov et al. (2022), who report the preservation of growth lines which were previously documented only in Silurian specimens.[44]

References

Matsukawa, M. (2022). "Barremian – Aptian ammonite biostratigraphy of the Sanchu Cretaceous, Japan". Cretaceous Research: Article 105245. doi:10.1016/j.cretres.2022.105245.
Rogov, M. A.; Lutikov, O. A. (2022). "A Dactylioceras – Meleagrinella (Clathrolima) assemblage from the Agardhbukta (eastern coast of Spitsbergen): a first in situ Toarcian molluscan occurrence from Svalbard providing interregional correlation". Norwegian Journal of Geology. doi:10.17850/njg102-1-2.
Korn, D.; Bockwinkel, J. (2022). "The tornoceratid ammonoids from the Roteisenstein Formation of Dillenburg (Cephalopoda, Ammonoidea)". European Journal of Taxonomy (806): 32–51. doi:10.5852/ejt.2022.806.1699.
Delanoy, G.; Baudouin, C.; Pictet, A.; Moreno-Bedmar, J.; Frau, C.; Matrion, B. (2022). "The genera Roloboceras Casey, 1954, and Megatyloceras Humphrey, 1949 (Ammonoidea, Ancyloceratina, Douvilleiceratidae), from the Lower Aptian of Ardèche (SE France) - Taxonomic and biostratigraphic implications". Carnets de Géologie. 22 (2): 7–109. doi:10.2110/carnets.2022.2202. S2CID 247397765.
Gelin, G. R. J.-P. (2022). "Nejdia Arkell, 1952, an Early Jurassic (Toarcian) ammonite from the Marrat formation, Central Saudi Arabia". Historical Biology: An International Journal of Paleobiology. Online edition: 1–17. doi:10.1080/08912963.2021.1999940. S2CID 247433723.
Ehiro, M. (2022). "Early Anisian (Aegean) ammonoids from the Fukkoshi Formation (Inai Group) with special reference to the Olenekian/Anisian boundary in the South Kitakami Belt, Northeast Japan" (PDF). Bulletin of the Tohoku University Museum. 21: 39–84.
Đaković, M.; Krystyn, L.; Sudar, M. (2022). "The Middle Smithian (Early Triassic) ammonoids of Gornji Brčeli (Southern Montenegro)". Rivista Italiana di Paleontologia e Stratigrafia. 128 (2): 411–430. doi:10.54103/2039-4942/12253.
Bischof, E. A.; Schlüter, N.; Lehmann, J. (2022). "Geometric morphometric analysis of morphologic disparity, intraspecific variation and ontogenetic allometry of beyrichitine ammonoids". PLOS ONE. 17 (2): e0263524. doi:10.1371/journal.pone.0263524. PMC 8830730. PMID 35143539.
Weber, C.; Hautmann, M.; Tajika, A.; Klug, C. (2022). "Is the relative thickness of ammonoid septa influenced by ocean acidification, phylogenetic relationships and palaeogeographic position?". Swiss Journal of Palaeontology. 141 (1): Article 4. doi:10.1186/s13358-022-00246-2.
Hebdon, N.; Ritterbush, K. A.; Choi, Y. J.; Peterman, D. (2022). "Reevaluating hydrodynamic performance of Late Triassic–Early Jurassic ammonoid shells with a 1D trajectory model". Geobios. in press. doi:10.1016/j.geobios.2022.02.002.
Kawakami, Y.; Uchiyama, N.; Wani, R. (2022). "Ontogenetic trajectories of septal spacing and conch shape in the Late Cretaceous gaudryceratid ammonoids: implications for their post-embryonic palaeoecology". Palaeontology. 65. doi:10.1111/pala.12587. S2CID 246140718.
Efremenko, V. D. (2022). "New belemnites of the family Cylindroteuthididae from the Lower Cretaceous of northern East Siberia, Nordvik Peninsula". Paleontological Journal. 56 (1): 37–46. doi:10.1134/S003103012201004X.
Niko, S.; Mapes, R. H.; Seuss, B. (2022). "Virgilian (Late Pennsylvanian) coiled nautiloids from the Finis Shale Member of the Graham Formation in Texas, southern Midcontinent North America" (PDF). Bulletin of the Tohoku University Museum. 21: 1–19.
Aubrechtová, M.; Korn, D. (2022). "Taxonomy and ontogeny of the Lituitida (Cephalopoda) from Orthoceratite Limestone erratics (Middle Ordovician)". European Journal of Taxonomy (799): 1–108. doi:10.5852/ejt.2022.799.1681. S2CID 247340759.
Korn, D.; Miao, L.; Bockwinkel, J. (2022). "The nautiloids from the Early Carboniferous Dalle à Merocanites of Timimoun, western Algeria". European Journal of Taxonomy (789): 104–129. doi:10.5852/ejt.2022.789.1635. S2CID 246398203.
Ehiro, M. (2022). "Middle Permian cephalopods of the Takakurayama Formation in the Yaguki area, the southwestern margin of the South Kitakami Belt, Northeast Japan" (PDF). Bulletin of the Tohoku University Museum. 21: 21–38.
Whalen, C. D.; Landman, N. H. (2022). "Fossil coleoid cephalopod from the Mississippian Bear Gulch Lagerstätte sheds light on early vampyropod evolution". Nature Communications. 13 (1): Article number 1107. doi:10.1038/s41467-022-28333-5. PMC 8904582. PMID 35260548.
Imbler, Sabrina (8 March 2022). "Fossil of Vampire Squid's Oldest Ancestor Is Named for Biden". The New York Times. Retrieved 8 March 2022.
Niko, S.; Ehiro, M. (2022). "Tohokubelus gen. nov., the Oldest Belemnite from the Olenekian (Lower Triassic) of Northeast Japan". Paleontological Research. 26 (2): 115–123. doi:10.2517/PR200036.
Pohle, A.; Kröger, B.; Warnock, R. C. M.; King, A. H.; Evans, D. H.; Aubrechtová, M.; Cichowolski, M.; Fang, X.; Klug, C. (2022). "Early cephalopod evolution clarified through Bayesian phylogenetic inference". BMC Biology. 20 (1): Article number 88. doi:10.1186/s12915-022-01284-5. PMC 9008929. PMID 35421982.
Lukeneder, P.; Lukeneder, A. (2022). "Mineralized belemnoid cephalic cartilage from the late Triassic Polzberg Konservat-Lagerstätte (Austria)". PLOS ONE. 17 (4): e0264595. doi:10.1371/journal.pone.0264595. PMC 9020720. PMID 35442996.
Goedert, J. L.; Kiel, S.; Tsai, C.-H. (2022). "Miocene Nautilus (Mollusca, Cephalopoda) from Taiwan, and a review of the Indo-Pacific fossil record of Nautilus". Island Arc. 31 (1): e12442. doi:10.1111/iar.12442.
Berezovsky, A. A. (2022). "Revision of Bathytormus (Crassatellidae, Bivalvia) of the Middle Eocene of Krivoi Rog (Ukraine)". Paleontological Journal. 56 (1): 24–36. doi:10.1134/S0031030122010038.
Berezovsky, A. A. (2022). "A new species of Crassatella (Bivalvia) from the Upper Eocene of central Ukraine". Paleontological Journal. 56 (2): 160–164. doi:10.1134/S0031030122020046.
Biakov, A. S. (2022). "New bivalves of the genus Myonia (Pholadomyida, Sanguinolitidae) from Permian deposits of northeast Russia". Paleontological Journal. 56 (2): 154–159. doi:10.1134/S003103012202006X.
Silantiev, V.; Marchetti, L.; Ronchi, A.; Schirolli, P.; Scholze, F.; Urazaeva, M. (2022). "Permian non-marine bivalves from the Collio and Guncina formations (Southern Alps, Italy): revised biostratigraphy and palaeobiogeography". Rivista Italiana di Paleontologia e Stratigrafia. 128 (1): 43–67. doi:10.54103/2039-4942/17050. S2CID 245899626.
Opazo, L. F.; Twitchett, R. J. (2022). "Bivalve body-size distribution through the Late Triassic mass extinction event". Paleobiology. in press: 1–26. doi:10.1017/pab.2021.38. S2CID 246311396.
Isaji, S.; Haga, T.; Kashiwagi, K. (2022). "Early Cretaceous Small-Sized Gastropods from the Shallow Marine Deposits of the Kimigahama Formation, Choshi Group, Japan". Paleontological Research. 26 (1): 31–54. doi:10.2517/PR20009. S2CID 245476443.
Pavia, G.; Dulai, A.; Festa, A.; Gennari, R.; Pavia, M.; Carnevale, G. (2022). "Palaeontology of the Upper Pliocene marine deposits of Rio Vaccaruzza, Villalvernia (Piedmont, NW Italy)". Rivista Italiana di Paleontologia e Stratigrafia. 128 (1): 129–210. doi:10.54103/2039-4942/15178 (inactive 2022-03-09).{{cite journal}}: CS1 maint: DOI inactive as of March 2022 (link)
Karapunar, B.; Nützel, A.; Seuss, B.; Mapes, R. H. (2022). "Taxonomy and diversity of slit-band gastropods (Order Pleurotomariida) and some slit bearing Caenogastropoda from the Pennsylvanian of the USA". Papers in Palaeontology. 8 (2): e1417. doi:10.1002/spp2.1417.
Harzhauser, M.; Landau, B.; Janssen, E. (2022). "The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera". Zootaxa. 5123 (1): 1–172. doi:10.11646/zootaxa.5123.1.1.
Gründel, R.; Hostettler, B.; Menkveld-Gfeller, U. (2022). "Gastropoden aus der Korallenrifffazies der St-Ursanne-Formation (mittleres Oxfordien) des Schweizer Jura. 3. Die Unterklasse Caenogastropoda Cox, 1960". Revue de Paléobiologie, Genève. 41 (1): 29–84. doi:10.5281/zenodo.6022757.
Psarras, C.; Merle, D.; Koskeridou, E. (2022). "Late Miocene Conidae (Mollusca: Gastropoda) of Crete (Greece). Part 2". European Journal of Taxonomy (816): 1–70. doi:10.5852/ejt.2022.816.1747.
Bail, P.; Ledon, D.; Pacaud, J.-M. (2022). "Description de deux espèces nouvelles de Lyria Gray, 1847 (Gastropoda : Volutidae) de l'Oligocène supérieur (Chattien) et du Miocène inférieur (Aquitanien) du sud-ouest de la France". Xenophora Taxonomy. 35: 3–7.
Raisossadat, S. N.; Guzhov, A.; Nazemi, R.; Motamedalshariati, M. (2022). "Gastropoda from Lower Cretaceous Deposits of Nimbolook Area, Eastern Iran". Stratigraphy and Geological Correlation. 30 (1): 65–74. doi:10.1134/S0869593822010063. S2CID 247413063.
Karapunar, B.; Nützel, A.; Ketwetsuriya, C. (2022). "A low-diversity Peruvispira-dominated gastropod assemblage from the Permian Ratburi Group of Central Thailand". Alcheringa: An Australasian Journal of Palaeontology. doi:10.1080/03115518.2022.2050814.
Liverani, V.; Wieneke, U. (2022). "Some more species in the genus Spinatus Dekkers et al., 2020 (Gastropoda, Strombidae)". Conchylia. 52 (3–4): 81–87.
Neubauer, T. A.; Hauffe, T.; Silvestro, D.; Scotese, C. R.; Stelbrink, B.; Albrecht, C.; Delicado, D.; Harzhauser, M.; Wilke, T. (2022). "Drivers of diversification in freshwater gastropods vary over deep time". Proceedings of the Royal Society B: Biological Sciences. 289 (1968): Article ID 20212057. doi:10.1098/rspb.2021.2057. PMC 8808086. PMID 35105242.
Neubauer, T. A.; Harzhauser, M. (2022). "Onset of Late Cretaceous diversification in Europe's freshwater gastropod fauna links to global climatic and biotic events". Scientific Reports. 12 (1): Article number 2684. doi:10.1038/s41598-022-06557-1. PMC 8854554. PMID 35177660.
Squires, R. L. (2022). "The earliest Ancistrolepis (Gastropoda: Buccinidae) and its geologic implications". PaleoBios. 39 (2): 1–11. doi:10.5070/P939257077.
Vendrasco, M. J.; Powell, C. L.; LaFollette, P. (2022). "The first Miocene chiton fauna from the northeastern Pacific". Journal of Paleontology: 1–14. doi:10.1017/jpa.2022.4.
Peel, J. S.; Kouchinsky, A. (2022). "Middle Cambrian (Miaolingian Series, Wuliuan Stage) molluscs and mollusc-like microfossils from North Greenland (Laurentia)". Bulletin of the Geological Society of Denmark. 70: 69–104. doi:10.37570/bgsd-2022-70-06.
Dell'Angelo, B.; Landau, B. M.; da Silva, C. M.; Sosso, M. (2022). "Biogeography of northeastern Atlantic Neogene chitons (Mollusca, Polyplacophora): New data from the Pliocene of Portugal". Journal of Paleontology. Online edition: 1–25. doi:10.1017/jpa.2022.1. S2CID 247329004.
Gubanov, A. P.; Bakayeva, S.; Ebbestad, J. O. R.; Bogolepova, O. K. (2022). "The Middle Ordovician Jinonicella (Mollusca) from Belarus and Ukraine". GFF. doi:10.1080/11035897.2022.2047773.

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[CRMatsukawa-1] Matsukawa, M. (2022). "Barremian – Aptian ammonite biostratigraphy of the Sanchu Cretaceous, Japan". Cretaceous Research: Article 105245. doi:10.1016/j.cretres.2022.105245.

[2] Rogov, M. A.; Lutikov, O. A. (2022). "A Dactylioceras – Meleagrinella (Clathrolima) assemblage from the Agardhbukta (eastern coast of Spitsbergen): a first in situ Toarcian molluscan occurrence from Svalbard providing interregional correlation". Norwegian Journal of Geology. doi:10.17850/njg102-1-2.

[EJT806-3] Korn, D.; Bockwinkel, J. (2022). "The tornoceratid ammonoids from the Roteisenstein Formation of Dillenburg (Cephalopoda, Ammonoidea)". European Journal of Taxonomy (806): 32–51. doi:10.5852/ejt.2022.806.1699.

[4] Delanoy, G.; Baudouin, C.; Pictet, A.; Moreno-Bedmar, J.; Frau, C.; Matrion, B. (2022). "The genera Roloboceras Casey, 1954, and Megatyloceras Humphrey, 1949 (Ammonoidea, Ancyloceratina, Douvilleiceratidae), from the Lower Aptian of Ardèche (SE France) - Taxonomic and biostratigraphic implications". Carnets de Géologie. 22 (2): 7–109. doi:10.2110/carnets.2022.2202. S2CID 247397765.

[5] Gelin, G. R. J.-P. (2022). "Nejdia Arkell, 1952, an Early Jurassic (Toarcian) ammonite from the Marrat formation, Central Saudi Arabia". Historical Biology: An International Journal of Paleobiology. Online edition: 1–17. doi:10.1080/08912963.2021.1999940. S2CID 247433723.

[Psilokhvalynites-6] Ehiro, M. (2022). "Early Anisian (Aegean) ammonoids from the Fukkoshi Formation (Inai Group) with special reference to the Olenekian/Anisian boundary in the South Kitakami Belt, Northeast Japan" (PDF). Bulletin of the Tohoku University Museum. 21: 39–84.

[7] Đaković, M.; Krystyn, L.; Sudar, M. (2022). "The Middle Smithian (Early Triassic) ammonoids of Gornji Brčeli (Southern Montenegro)". Rivista Italiana di Paleontologia e Stratigrafia. 128 (2): 411–430. doi:10.54103/2039-4942/12253.

[8] Bischof, E. A.; Schlüter, N.; Lehmann, J. (2022). "Geometric morphometric analysis of morphologic disparity, intraspecific variation and ontogenetic allometry of beyrichitine ammonoids". PLOS ONE. 17 (2): e0263524. doi:10.1371/journal.pone.0263524. PMC 8830730. PMID 35143539.

[9] Weber, C.; Hautmann, M.; Tajika, A.; Klug, C. (2022). "Is the relative thickness of ammonoid septa influenced by ocean acidification, phylogenetic relationships and palaeogeographic position?". Swiss Journal of Palaeontology. 141 (1): Article 4. doi:10.1186/s13358-022-00246-2.

[10] Hebdon, N.; Ritterbush, K. A.; Choi, Y. J.; Peterman, D. (2022). "Reevaluating hydrodynamic performance of Late Triassic–Early Jurassic ammonoid shells with a 1D trajectory model". Geobios. in press. doi:10.1016/j.geobios.2022.02.002.

[11] Kawakami, Y.; Uchiyama, N.; Wani, R. (2022). "Ontogenetic trajectories of septal spacing and conch shape in the Late Cretaceous gaudryceratid ammonoids: implications for their post-embryonic palaeoecology". Palaeontology. 65. doi:10.1111/pala.12587. S2CID 246140718.

[PJ561-12] Efremenko, V. D. (2022). "New belemnites of the family Cylindroteuthididae from the Lower Cretaceous of northern East Siberia, Nordvik Peninsula". Paleontological Journal. 56 (1): 37–46. doi:10.1134/S003103012201004X.

[NikoMapesSeuss-13] Niko, S.; Mapes, R. H.; Seuss, B. (2022). "Virgilian (Late Pennsylvanian) coiled nautiloids from the Finis Shale Member of the Graham Formation in Texas, southern Midcontinent North America" (PDF). Bulletin of the Tohoku University Museum. 21: 1–19.

[EJT799-14] Aubrechtová, M.; Korn, D. (2022). "Taxonomy and ontogeny of the Lituitida (Cephalopoda) from Orthoceratite Limestone erratics (Middle Ordovician)". European Journal of Taxonomy (799): 1–108. doi:10.5852/ejt.2022.799.1681. S2CID 247340759.

[EJT789-15] Korn, D.; Miao, L.; Bockwinkel, J. (2022). "The nautiloids from the Early Carboniferous Dalle à Merocanites of Timimoun, western Algeria". European Journal of Taxonomy (789): 104–129. doi:10.5852/ejt.2022.789.1635. S2CID 246398203.

[16] Ehiro, M. (2022). "Middle Permian cephalopods of the Takakurayama Formation in the Yaguki area, the southwestern margin of the South Kitakami Belt, Northeast Japan" (PDF). Bulletin of the Tohoku University Museum. 21: 21–38.

[17] Whalen, C. D.; Landman, N. H. (2022). "Fossil coleoid cephalopod from the Mississippian Bear Gulch Lagerstätte sheds light on early vampyropod evolution". Nature Communications. 13 (1): Article number 1107. doi:10.1038/s41467-022-28333-5. PMC 8904582. PMID 35260548.

[18] Imbler, Sabrina (8 March 2022). "Fossil of Vampire Squid's Oldest Ancestor Is Named for Biden". The New York Times. Retrieved 8 March 2022.

[19] Niko, S.; Ehiro, M. (2022). "Tohokubelus gen. nov., the Oldest Belemnite from the Olenekian (Lower Triassic) of Northeast Japan". Paleontological Research. 26 (2): 115–123. doi:10.2517/PR200036.

[20] Pohle, A.; Kröger, B.; Warnock, R. C. M.; King, A. H.; Evans, D. H.; Aubrechtová, M.; Cichowolski, M.; Fang, X.; Klug, C. (2022). "Early cephalopod evolution clarified through Bayesian phylogenetic inference". BMC Biology. 20 (1): Article number 88. doi:10.1186/s12915-022-01284-5. PMC 9008929. PMID 35421982.

[21] Lukeneder, P.; Lukeneder, A. (2022). "Mineralized belemnoid cephalic cartilage from the late Triassic Polzberg Konservat-Lagerstätte (Austria)". PLOS ONE. 17 (4): e0264595. doi:10.1371/journal.pone.0264595. PMC 9020720. PMID 35442996.

[22] Goedert, J. L.; Kiel, S.; Tsai, C.-H. (2022). "Miocene Nautilus (Mollusca, Cephalopoda) from Taiwan, and a review of the Indo-Pacific fossil record of Nautilus". Island Arc. 31 (1): e12442. doi:10.1111/iar.12442.

[23] Berezovsky, A. A. (2022). "Revision of Bathytormus (Crassatellidae, Bivalvia) of the Middle Eocene of Krivoi Rog (Ukraine)". Paleontological Journal. 56 (1): 24–36. doi:10.1134/S0031030122010038.

[24] Berezovsky, A. A. (2022). "A new species of Crassatella (Bivalvia) from the Upper Eocene of central Ukraine". Paleontological Journal. 56 (2): 160–164. doi:10.1134/S0031030122020046.

[PJ562Myonia-25] Biakov, A. S. (2022). "New bivalves of the genus Myonia (Pholadomyida, Sanguinolitidae) from Permian deposits of northeast Russia". Paleontological Journal. 56 (2): 154–159. doi:10.1134/S003103012202006X.

[RIPSPalaeomutela-26] Silantiev, V.; Marchetti, L.; Ronchi, A.; Schirolli, P.; Scholze, F.; Urazaeva, M. (2022). "Permian non-marine bivalves from the Collio and Guncina formations (Southern Alps, Italy): revised biostratigraphy and palaeobiogeography". Rivista Italiana di Paleontologia e Stratigrafia. 128 (1): 43–67. doi:10.54103/2039-4942/17050. S2CID 245899626.

[27] Opazo, L. F.; Twitchett, R. J. (2022). "Bivalve body-size distribution through the Late Triassic mass extinction event". Paleobiology. in press: 1–26. doi:10.1017/pab.2021.38. S2CID 246311396.

[Choshipleura-28] Isaji, S.; Haga, T.; Kashiwagi, K. (2022). "Early Cretaceous Small-Sized Gastropods from the Shallow Marine Deposits of the Kimigahama Formation, Choshi Group, Japan". Paleontological Research. 26 (1): 31–54. doi:10.2517/PR20009. S2CID 245476443.

[RIPSRioVaccaruzza-29] Pavia, G.; Dulai, A.; Festa, A.; Gennari, R.; Pavia, M.; Carnevale, G. (2022). "Palaeontology of the Upper Pliocene marine deposits of Rio Vaccaruzza, Villalvernia (Piedmont, NW Italy)". Rivista Italiana di Paleontologia e Stratigrafia. 128 (1): 129–210. doi:10.54103/2039-4942/15178 (inactive 2022-03-09).{{cite journal}}: CS1 maint: DOI inactive as of March 2022 (link)

[Eirlysella-30] Karapunar, B.; Nützel, A.; Seuss, B.; Mapes, R. H. (2022). "Taxonomy and diversity of slit-band gastropods (Order Pleurotomariida) and some slit bearing Caenogastropoda from the Pennsylvanian of the USA". Papers in Palaeontology. 8 (2): e1417. doi:10.1002/spp2.1417.

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