2022 in paleomammalogy

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This paleomammology list records new fossil mammal taxa that were described during the year 2022, as well as notes other significant paleomammalogy discoveries and events which occurred during 2022.

Afrotherians

Proboscidea

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Platybelodon tetralophus[1]	Sp. nov	In press	Wang & Li	Miocene	Tunggur Formation	China

Proboscidean research

Partial skeleton of a specimen of "Mammut" borsoni, representing one of the most recent record of mammutids in Europe reported to date, is described from the Villafranchian of Kaltensundheim (Thuringia, Germany) by Koenigswald et al. (2022).[2]
A study on the carbon and oxygen isotope ratios in teeth of a sub-adult mastodon found in southern Brazil is published by Lopes et al. (2022), who interpret their findings as indicative of a diet shift during the life of the animal, and indicating that mastodons were able to change their diets at shorter timescales than can be addressed from the analysis of isolated teeth.[3]
Fossil material of a member or a relative of the genus Sinomastodon is described from the Quaternary of the Kashmir Valley by Parray et al. (2022), representing the youngest record of a gomphothere from the Indian Subcontinent reported to date.[4]
A study on the origin, dispersal and ecology of gomphotheres in South America is published by Alberdi & Prado (2022).[5]
A study on the range of size variation in palaeoloxodont elephants from Sicily, Favignana and Malta, inhabiting the Siculo-Maltese Palaeoarchipelago during the Pleistocene, and on possible reasons for size differences of these elephants is published by Scarborough (2022).[6]

Sirenia

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Dakhlasiren[7]	Gen. et sp. nov	In press	Zouhri, Zalmout & Gingerich	Late Eocene	Samlat Formation	Western Sahara	A member of the family Protosirenidae. The type species is D. marocensis.

Sirenian research

Description of the anatomy of the skull of Sobrarbesiren cardieli and a study on the affinities of this taxon is published by Díaz-Berenguer et al. (2022).[8]

Other afrotherians

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images

Euarchontoglires

Primates

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Chlorocebus ngedere[9]	Sp. nov		Arenson et al.	Early Pleistocene	Lower Ngaloba Beds	Tanzania	A guenon
Sawecolobus[10]	Gen. et sp. nov	Valid	Gommery et al.	Late Miocene	Lukeino Formation	Kenya	An Old World monkey belonging to the subfamily Colobinae. The type species is S. lukeinoensis.

Primate research

A study on the talar and calcaneal morphology in Eocene primates from the Vastan lignite mine (Gujarat, India), and on its implicatios for the knowledge of the locomotor capabilities of these primates, is published by Llera Martín, Rose & Sylvester (2022).[11]
A study on the internal nasal anatomy of Homunculus patagonicus, and on its implications for the knowledge of the phylogenetic affinities of this monkey, is published by Lundeen & Kay (2022).[12]
A study on the phylogenetic relationships for Middle-Late Miocene fossil apes is published by Pugh (2022).[13]
Description of new fossil material of apes from the Miocene locality of Berg Aukas (Namibia) and new information on the locality of the ape mandible from the Miocene of Niger described by Pickford et al. (2008)[14] is published by Mocke et al. (2022), who evaluate the implications of these fossils for the knowledge of the evolution of the African apes.[15]
A study on the occurrence and morphology of calcar femorale in extant and fossil hominids is published by Cazenave et al. (2022), who interpret their findings as indicating that this structure cannot be considered as a diagnostic feature of habitual bipedal locomotion.[16]
Dental remains of Gigantopithecus blacki, possibly belonging to one of the latest relict populations of Gigantopithecus, are described from the Upper Pleistocene deposits of the Lang Trang cave (Vietnam) by Lopatin, Maschenko & Dac (2022).[17]
Atypical tooth wear, similar to tooth wear previously reported in fossil hominins and regarded as possible evidence of early cultural habits, is reported in a sample of extant Japanese macaques from Koshima Island by Towle et al. (2022), who interpret the atypical wear patterns as likely caused by accidental ingestion of sand and oral processing of marine mollusks, and evaluate the impications of this finding for interpretations of similar wear in fossil hominins.[18]
A review aiming to determine the value of extant primates as models for reconstructions of fossil hominin stone tool culture is published by Bandini, Harrison & Motes-Rodrigo (2022).[19]

General paleoanthropology

A study on the likely diet of members of the genus Paranthropus is published by Sponheimer et al. (2022).[20]
A study on the origins of the complex birth pattern characteristic of modern humans, based on data from simulations of the birth process in australopithecines, is published by Frémondière et al. (2022).[21]
A study on the habitat types at the Woranso-Mille site (Ethiopia) during the Pliocene, and on factors which allowed the coexistence of more than one species of Australopithecus at the site, is published by Denise Su & Yohannes Haile-Selassie (2022).[22]
A study comparing the distal portion of the fibula of Australopithecus afarensis and extant humans and apes, aiming to determine the correlates of distal fibular shape with arboreal behavior in extant hominids and fossil hominins is published by Marchi et al. (2022).[23]
A calcaneus of an early hominin, with a morphology that is intermediate between humans and nonhuman apes, is described from the Kromdraai fossil site (South Africa) by Harper et al. (2022).[24]
A study on the impact of climate variability on the evolution of early African Homo, Eurasian Homo erectus, Homo heidelbergensis, Neanderthals and modern humans is published by Timmermann et al. (2022).[25]
A vertebra of a juvenile hominin is described from the early Pleistocene site of 'Ubeidiya (Israel) by Barash et al. (2022), who estimate the adult size of this hominin as comparable to early Pleistocene large-bodied hominins from Africa, and interpret this finding as the earliest large-bodied hominin remains from the Levantine corridor reported to date, distinct from other early Eurasian hominins, sharing affinities to East African large-bodied hominins, and supporting the occurrence of several Pleistocene dispersals of hominins out of Africa.[26]
A study on the 2.6 to 1.2 million years old zooarchaeological record of eastern Africa, aiming to determine whether the zooarchaeological record preserves sustained increase in the amount of evidence for hominin carnivory after the appearance of Homo erectus, is published by Barr et al. (2022).[27]
A study on the lesions of Dmanisi skull D2280 is published by Margvelashvili et al. (2022), who interpret the studied pathologies as evidence of blunt force trauma possibly caused by interpersonal violence, as well as evidence of treponemal disease.[28]
Evidence from the Zhoukoudian Locality 1 interpreted as indicative of controlled use of fire by Peking Man is presented by Huang, Li & Gao (2022).[29]
Description of the cochlear morphology of two individuals of Homo erectus from the Indonesian site Sangiran (Sangiran 2 and 4), comparing them with a sample australopiths and Middle to Late Pleistocene and extant humans, is published by Urciuoli et al. (2022).[30]
A study on the morphological variability among Middle Pleistocene Chinese hominins, aiming to determine the evolutionary processes that shaped hominin variation in eastern Eurasia during the Middle Pleistocene, is published by Liu et al. (2022).[31]
A study on the external and internal tooth structure in Homo luzonensis, and on its implications for the knowledge of the affinities of this species, is published by Zanolli et al. (2022).[32]
The first reconstruction of a fairly complete hominin posterior cranium from the late Middle Pleistocene Xujiayao site (China), and a study on the endocranial capacity of this cranium, is published by Wu et al. (2022), who interpret this specimen as the earliest evidence of a brain size that falls in the upper range of Neanderthals and modern Homo sapiens, and evaluate its implications for the knowledge of the evolution of the hominin brain size.[33]
A study on the Late Pleistocene human population dynamics, aiming to determine how the process of the replacement of Eurasian archaic humans by anatomically modern human populations dispersing from Africa unfolded, is published by Vahdati et al. (2022).[34]
A study on the development of teeth in Pleistocene hominins from the Gran Dolina and the Sima de los Huesos sites of the Sierra de Atapuerca (Spain) is published by Modesto-Mata et al. (2022).[35]
A study on the taphonomic features of the hominin skull remains from the Sima de los Huesos sample, aiming to create a catalog of modifications to crania and mandibles (including antemortem, perimortem and postmortem skeletal disturbances) within this sample, is published by Sala et al. (2022).[36]
A study on the impact of the sexual dimorphism, ancestry and lifestyle effects on lordosis in a large sample of modern humans and Neanderthals is published by Williams et al. (2022), who interpret their findings as casting doubt on proposed locomotor and postural differences between modern humans and Neanderthals based on inferred lumbar lordosis (or lack thereof), and indicating that future studies should not compare remains of fossil hominins and preindustrial modern humans to samples from sedentary, industrialized populations, but rather to the remains of individuals that engaged in more active, traditional lifestyles.[37]
Four teeth of Neanderthals, belonging to at least two individuals (an adult and a child) and representing the earliest evidence of Neanderthal spread into the Eastern Mediterranean Area reported to date, are described from the Chibanian of the Velika Balanica cave (Serbia) by Roksandic et al. (2022).[38]
A study on the age of the Omo remains is published by Vidal et al. (2022).[39]
A study on the anatomy of the brain, braincase and bony labyrinth of the Border Cave 1 cranium is published by Beaudet et al. (2022).[40]
Reconstruction of the eastern African environments inhabited inhabited by early human populations during the Middle Stone Age, evaluating the role of shifting environmental conditions on the distribution and variability of dated Middle Stone Age assemblages, is published by Timbrell et al. (2022).[41]
Evidence of four periods of human occupation between c. 210,000 and 120,000 years ago is reported from Jebel Faya (United Arab Emirates) by Bretzke et al. (2022), who evaluate the implications of these findings for the knowledge of the impact of arid conditions on Paleolithic human populations in Arabia.[42]
Possible evidence of use of fruits and wood from olive trees by the early Homo sapiens around 100,000 years ago is reported from Morocco by Marquer et al. (2022).[43]
Evidence of the production of ostrich eggshell artefacts, long-distance transportation of marine molluscs and systematic use of heat shatter in stone tool production approximately 92–80 thousand years before the present is reported from the Varsche Rivier 003 site (South Africa) by Mackay et al. (2022), who evaluate the implications of these findings for the knowledge of the processes of innovation and cultural transmission in southern Africa during the Middle Stone Age.[44]
Hominin fossils interpreted as evidence of the earliest known arrival of modern humans in Europe (between 56,800 and 51,700 calibrated years before the present) are described from the Grotte Mandrin (France) by Slimak et al. (2022).[45]
A study on the microstructure and likely origin of the material used to produce the Venus of Willendorf is published by Weber et al. (2022).[46]
The earliest ochre-processing feature in Eastern Asia reported to date, a bone tool and a distinctive miniaturized lithic assemblage with bladelet-like tools bearing traces of hafting, representing a cultural assembly of traits that is unique for Eastern Asia, is described from the approximately 40,000-year-old Xiamabei site (China) by Wang et al. (2022).[47]
A study on patterns in the stratigraphic integrity of early North American archeological sites, and on their implications for the knowledge of the timing of human arrival to North America, is published by Surovell et al. (2022).[48]
A study on the authenticity of the potential Ice Age rock art of Serranía de la Lindosa (Colombia) is published by Iriarte et al. (2022), who argue that there are sound grounds to consider the studied paintings as ancient and likely representing now-extinct Ice Age megafauna.[49]
Lipson et al. (2022) present new genome-wide ancient DNA data from three Late Pleistocene and three early to middle Holocene individuals associated with Late Stone Age technologies from Kisese II and Mlambalasi Rockshelters in Tanzania, Fingira and Hora 1 Rockshelters in Malawi and Kalemba Rockshelter in Zambia, and study changes in regional- and continental-scale population structures in sub-Saharan Africa during the Late Pleistocene and early Holocene.[50]
Computational biologists report the largest detailed human genetic genealogy, unifying human genomes from many sources for insights about human history, ancestry and evolution. It demonstrates a novel computational method for estimating how human DNA is related, in specific as a series of 13 million linked trees along the genome, a tree-sequence, which has also been called "the largest human family tree".[51][52][53]

Rodentia

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Hydrochoerus hesperotiganites[54]	Sp. nov	Valid	White et al.	Late Pleistocene		United States ( California)	A species of Capybara.	Hydrochoerus hesperotiganites in dorsal view
Mus kerati[55]	Sp. nov	Valid	Stoetzel & Pickford	Middle Pleistocene		Algeria	A species of Mus.
Paraethomys geraadsi[55]	Sp. nov	Valid	Stoetzel & Pickford	Middle Pleistocene		Algeria	A member of the family Muridae belonging to the subfamily Murinae.
Pararhizomys parvulus[56]	Sp. nov	In press	Wang	Miocene	Probably Liushu Formation	China	A member of the family Spalacidae belonging to the subfamily Tachyoryctoidinae and the tribe Pararhizomyini.
Shapajamys minor[57]	Sp. nov	In press	Arnal et al.	Paleogene	Santa Rosa fossil site	Peru	A caviomorph rodent.
Tobienia fejfari[58]	Sp. nov	In press	Tesakov & Bondarev	Late Pliocene		Russia	A member of the tribe Lemmini.
Ucayalimys amahuacensis[57]	Sp. nov	In press	Arnal et al.	Paleogene	Santa Rosa fossil site	Peru	A caviomorph rodent, possibly a member of Chinchilloidea.
Vucetichimys[57]	Gen. et sp. nov	In press	Arnal et al.	Paleogene	Santa Rosa fossil site	Peru	A caviomorph rodent. The type species is V. pretrilophodoncia.

Rodent research

Description of new fossil material of Hystrix makapanensis from Olduvai Gorge (Tanzania) and a review of the African record of this species is published by Azzarà et al. (2022).[59]
The first description of the postcranial remains of Bathyergoides neotertiarius from the Miocene of Namibia is published by Bento Da Costa & Senut (2022), who evaluate the implications of the studied fossils for the knowledge of the behaviour of this rodent.[60]
Description of new fossil material and a study on the taxonomic diversity of dinomyids from the late Miocene-early Pliocene Cerro Azul Formation (Argentina) is published by Sostillo et al. (2022).[61]
A study on the validity of the genus Gyriabrus, and a revision of the species assigned to this genus, is published by Rasia (2022).[62]
Description of cavioid, chinchilloid and erethizontoid rodents from the Miocene Pampa Castillo fauna (Chile) and a study on their biochronologic and paleoenvironmental implications is published by McGrath et al. (2022).[63]
A study on the phylogenetic relationships of Paronychomys and Basirepomys is published by Kelly & Martin (2022).[64]
A study on the anatomy of the skull of Hispanomys moralesi is published by Carro-Rodríguez et al. (2022).[65]
Description of the anatomy of the holotype specimen of the Tenerife giant rat is published by Casanovas-Vilar & Luján (2022).[66]

Other euarchontoglires

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Pronolagus humpatensis[67]	Sp. nov	Valid	Sen & Pickford	Early Pleistocene		Angola	A red rock hare.

Laurasiatherians

Artiodactyla

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Afrotragulus akhtari[68]	Sp. nov	In press	Sánchez et al.	Miocene		Pakistan	A chevrotain.
Afrotragulus megalomilos[68]	Sp. nov	In press	Sánchez et al.	Miocene		Pakistan	A chevrotain.
Afrotragulus moralesi[68]	Sp. nov	In press	Sánchez et al.	Miocene		Pakistan	A chevrotain.
Archaebalaenoptera eusebioi[69]	Sp. nov	Valid	Bisconti et al.	Miocene (Tortonian)	Pisco Formation	Peru	A rorqual.
Ebusia[70]	Gen. et sp. nov	Valid	Moyà-Solà, Quintana Cardona & Köhler	Probably Neogene		Spain	A caprine bovid. The type species is E. moralesi.
Hemiauchenia mirim[71]	Sp. nov		Greco et al.	Late Pleistocene		Brazil	A camelid.
Listriodon dukkar[72]	Sp. nov	Valid	Van der Made et al.	Late Miocene		India	A member of the family Suidae belonging to the subfamily Listriodontinae.
Miophyseter[73]	Gen. et sp. nov	Valid	Kimura & Hasegawa	Miocene (Burdigalian)	Toyohama Formation	Japan	A Physeteroidea sperm whale. Type species M. chitaensis.
Neochorlakkia[74]	Gen. et sp. nov	Valid	Ducrocq et al.	Eocene	Pondaung Formation	Myanmar	A member of the family Dichobunidae. The type species is N. myaingensis.
Rododelphis[75]	Gen. et sp. nov	Valid	Bianucci et al.	Early Pleistocene	Lindos Formation	Greece	An oceanic dolphin belonging to the subfamily Globicephalinae. Type species is R. stamatiadisi.
Rusingameryx[76]	Gen. et comb. nov	In press	Pickford	Miocene		Kenya Namibia Uganda	An anthracothere. The type species is "Brachyodus" aequatorialis MacInnes (1951).

Artiodactyl research

A study on the diet of Hemiauchenia paradoxa, guanaco and vicuña from the Pleistocene of southern Brazil is published by Carrasco et al. (2022).[77]
Description of camel remains from the Tsagaan Agui Cave and the Tugrug Shireet open-air site (Mongolia), including fossil material of Camelus knoblochi, is published by Klementiev et al. (2022), who interpret their findings as evidence of survival of C. knoblochi in the Gobi Desert until the Last Glacial Maximum.[78]
A study on the relationship between functional occlusal traits, dental wear and increase in crown length in the third molars of Pliocene and Pleistocene African suids, aiming to determine the evolutionary trends of the functional occlusal traits in these suids in the context of their dietary ecology and potential selective pressures, is published by Yang et al. (2022).[79]
Review of the large-sized members of the genus Palaeotragus from the Vallesian of northern Greece, and a systematic revision of large-sized Late Miocene Eurasian members of the genus Palaeotragus, is published by Laskos & Kostopoulos (2022).[80]
New fossil material of a member of the genus Acteocemas belonging or related to the species A. infans, providing evidence that protoantlers of Acteocemas were able to be cast and re-grown (but also indicating that the lifespan of these protoantlers could be longer than that of antlers of modern deers, preventing them from assuming a similar cycle), is described from the Miocene site of Sant Andreu de la Barca (Spain) by Azanza et al. (2022).[81]
A study on the histology of ribs of Candiacervus, and on its implications for the knowledge of the longevity of this deer, is published by Miszkiewicz & Van Der Geer (2022).[82]
A study on the evolutionary history of the Siberian roe deers, as indicated by data from four ancient mitochondrial genomes generated from roe deer fossil specimens from northeastern China, is published by Deng et al. (2022).[83]
Description of new fossil material of Qurliqnoria cheni from the northern Tibetan Plateau, providing new information on the anatomy of this bovid, is published by Tseng et al. (2022), who evaluate the implications of this finding for the knowledge of the evolution of the Tibetan antelope.[84]
A relatively complete cranium and mandible of Brachyodus onoideus, providing new information on the anatomy of this anthracothere, is described by Pickford & MacLaren (2022).[85]
A study comparing changes in the skull anatomy during the ontogeny in Hippopotamus gorgops and extant hippopotamus, based on data from the skull of a juvenile specimen of H. gorgops from the Early Pleistocene site of Buia (Eritrea), is published by Martínez-Navarro et al. (2022).[86]
Fossil material of a basilosaurid cetacean is described from the Eocene Beloglinskaya Formation (Krasnodar Krai, Russia) by Tarasenko (2022), representing the first record of a basilosaurid in the studied region.[87]
Description of a new specimen of an archaic dolphin (belonging or related to the species Prosqualodon davidis) from the Miocene Gee Greensand (New Zealand), and a study on the implications of this specimen for the knowledge of the evolution of the brain of toothed whales, is published by Tanaka, Ortega & Fordyce (2022).[88]
The second specimen of Casatia thermophila, providing new information on the anatomy of this monodontid, is described from the Pliocene locality of Arcille (Italy) by Merella et al. (2022).[89]
Review of the fundamental morphological transformations that occurred at the origin stage of the baleen whales is published by Bisconti & Carnevale (2022).[90]

Carnivorans

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes
Adeilosmilus[91]	Gen. et comb. nov	In press	Jiangzuo, Werdelin & Sun	Miocene		Chad	A member of Machairodontinae; a new genus for "Machairodus" kabir Peigné et al. (2005).
Aragonictis[92]	Gen. et sp. nov	In press	Valenciano et al.	Miocene		Spain	A small-sized mustelid. The type species is A. araid.
Hadrokirus novotini[93]	Sp. nov	In press	Hafed et al.	Neogene		United States ( North Carolina)	An earless seal belonging to the subfamily Monachinae.
Homiphoca murfreesi[93]	Sp. nov	In press	Hafed et al.	Neogene		United States ( North Carolina)	An earless seal belonging to the subfamily Monachinae.
Longchuansmilus[94]	Gen. et sp. nov	In press	Jiangzuo et al.	Late Miocene		China	A member of Machairodontinae. The type species is L. xingyongi.
Mogharacyon[95]	Gen. et comb. nov	Valid	Morales & Pickford	Early Miocene		Egypt	A bear dog; a new genus for "Cynelos" anubisi Morlo et al. (2019).
Moralesictis[96]	Gen. et sp. nov	In press	Valenciano & Baskin	Hemphillian		United States ( Texas)	A mellivorine mustelid. The type species is M. intrepidus.
Namibiocyon[95]	Gen. et comb. nov	Valid	Morales & Pickford	Early Miocene		Namibia	A bear dog; a new genus for "Ysengrinia" ginsburgi Morales et al. (1998).
Panthera gombaszoegensis jinpuensis[97]	Ssp. nov	In press	Jiangzuo et al.	Middle Pleistocene		China
Panthera uncia pyrenaica[98]	Ssp. nov	Valid	Hemmer	Pleistocene (Marine Isotope Stage 14)		France	A subspecies of the snow leopard.
Percrocuta xixiaensis[99]	Sp. nov	Valid	Xiong	Miocene	Zhang’enbao Formation	China
Protocyon orocualensis[100]	Sp. nov	Valid	Ruiz-Ramoni, Wang & Rincón	Late Pliocene/Early Pleistocene		Venezuela	A Cerdocyonina canine. Announced in 2021; the final article version was published in 2022.
Taowu[91]	Gen. et sp. nov	In press	Jiangzuo, Werdelin & Sun	Early Pleistocene		China	A member of Machairodontinae. The type species is T. liui.

Carnivoran research

A study on the fossils of carnivorans from the Miocene (Messinian) of Cava Monticino (Italy), including fossil material of Eucyon monticinensis representing one of the oldest, certain records of the genus Eucyon in the Old World and fossil material of Mellivora benfieldi representing the northernmost record of the species and the only certain record of the genus Mellivora outside of Africa, is published by Bartolini-Lucenti, Madurell-Malapeira & Rook (2022).[101]
Revision of the carnivoran fauna from Libakos in the Pliocene-Pleistocene Grevena–Neapolis Basin (Greece), including the first record of the mustelid Pannonictis nestii from Greece, and a study on the age of this fauna is published by Koufos & Tamvakis (2022).[102]
Fossil material of Amphicyon giganteus is described from a travertine above a layer dated to MN7/8 in the Karacalar Silver Travertine Quarry (Gebeceler Formation, Turkey) by van der Hoek et al. (2022), representing the youngest record of this species reported to date.[103]
Description of new fossil material of Xenocyon lycaonoides from the Jinyuan Cave (China), confirming the presence of this species in eastern Asia during the early Middle Pleistocene, and a study on the affinities of this species is published by Jiangzuo et al. (2022).[104]
Description of a robust canid dentary from the Pliocene Glenns Ferry Formation (Hagerman Fossil Beds National Monument; Idaho, United States), and a study on the affinities of this specimen and on the diversity of Pliocene canids from Hagerman, is published by Prassack & Walkup (2022).[105]
Description of a wolf skull from Ponte Galeria (Rome, Italy), representing the first reliable occurrence of this taxon in Europe and the largest skull of a Middle Pleistocene canid from Europe known to date, is published by Iurino et al. (2022), who evaluate the implications of this specimen for the knowledge of the turnover between Canis mosbachensis and modern wolves.[106]
A study on the functional morphology of the skull of the Pleistocene badger Meles dimitrius is published by Savvidou et al. (2022).[107]
A study on the skeletal morphology, affinities and likely paleoecology of small-sized cave bears (originally assigned to the taxon Ursus savini) from the Imanay Cave (Russia) is published by Gimranov et al. (2022).[108]
A study on the microwear of the non-occlusal surface of incisors of the small cave bear and Ural cave bear from the Pleistocene of the Middle and South Urals, and on its implications for the knowledge of the trophic specialization of these cave bears, is published by Gimranov, Zykov & Kosintsev (2022).[109]
Review of the knowledge of the taxonomy and phylogeny, biology, distribution, occurrence and extinction times, and interaction with humans of large and small cave bears in the Urals is published by Gimranov & Kosintsev (2022).[110]
A study on the evolutionary history and phylogeography of ancient and modern brown bears, based on data from mitochondrial genomes of four ancient (~4.5–40 thousand years old) bears from South Siberia and modern bears from South Siberia and the Russian Far East, is published by Molodtseva et al. (2022).[111]
Fossil material of members of the genus Palaeogale is described from the Oligocene John Day Formation (Oregon, United States) by Famoso & Orcutt (2022), representing the first known records of this genus from the Pacific Northwest of North America.[112]
A well-preserved skull of Stenoplesictis minor is described from the Oligocene Quercy Phosphorites Formation (France) by de Bonis et al. (2022), who present a reconstructon of brain endocast, stapes and bony labyrinths of this specimen.[113]
A mandible of the largest specimen belonging to the genus Pachycrocuta reported to date, with dental morphology similar to that of Pachycrocuta from Zhoukoudian, is described from the Middle Pleistocene loess in Luoning (Henan, China) by Jiangzuo et al. (2022).[114]
Review of the fossil record and a revision of the species-level taxonomy of the genus Crocuta is published by Lewis & Werdelin (2022).[115]
A study on the fossil record of members of the genus Amphimachairodus in the Chinese Baode strata is published by Wang, Carranza-Castañeda & Tseng (2022), who interpret this record as evidence of anagenetic evolution of increasing size, and study the evolution of members of the genus Amphimachairodus on the basis of all Holarctic records.[116]
Revised reconstruction of the soft tissue and life appearance of Homotherium latidens is proposed by Antón et al. (2022).[117]
A study on feeding damage from Xenosmilus hodsonae in the large mammalian fauna from the Irvingtonian paleo-sinkhole Haile 21A (Florida, United States), and on its implications for the knowledge of the carcass processing capabilities of Xenosmilus and of the sabertooth paleoecology in the Pleistocene, is published by Domínguez-Rodrigo et al. (2022).[118]
New fossil material of a lynx belonging or related to the species Lynx issiodorensis is described from the Villafranchian site of La Puebla de Valverde (Spain) by Cuccu et al. (2022), who evaluate the implications of this finding for the knowledge of the European lynx fossil record.[119]
Description of Late Pleistocene remains of the Iberian lynx from Avenc del Marge del Moro (Garraf Massif, Catalonia, Spain) is published by Tura-Poch et al. (2022).[120]
Large felid remains assigned to the species Panthera fossilis are described from the Grotte de la Carrière in Eastern Pyrenees by Prat-Vericat et al. (2022), who evaluate the implications of these fossils for the knowledge of the paleobiology of P. fossilis.[121]
Two specimens of Panthera spelaea are described from the Middle and Late Pleistocene Songhua River fossil assemblages (China) by Sherani, Perng & Sherani (2022), representing the first records of this species from the Mammuthus-Coelodonta fauna from the Pleistocene assemblages of the Songhua River reported to date.[122]

Chiroptera

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images

Eulipotyphla

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Dzavui[123]	Gen. et sp. nov	Valid	Crespo & Jiménez-Hidalgo in Jiménez-Hidalgo, Guerrero-Arenas & Crespo	Oligocene (Rupelian)		Mexico	A gymnure. The type species is D. landeri.

Eulipotyphlean research

Fossil material of the erinaceid Galerix rutlandae and a talpid belonging to the subfamily Uropsilinae, representing the first known record of these families from the Miocene Siwalik exposures of India and the first record of an uropsiline from the Indian subcontinent, is described by Parmar, Norboo & Magotra (2022).[124]
Fossil material of Van Sung's shrew and Chodsigoa hoffmanni is described from the Pleistocene of the Tham Hai cave and Lang Trang cave (Vietnam) by Lopatin (2022), representing the first fossil records of these species and the first fossil remains of members of the genus Chodsigoa found outside China.[125]

Perissodactyla

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images

Perissodactyl research

A study on the evolutionary variation of shape in hindlimb long bones of members of Rhinocerotoidea, and on its relationship with mass, size and gracility, is published by Mallet et al. (2022).[126]
Description of new fossil material of Pliorhinus megarhinus from the early Pliocene of the Vera Basin (Spain) and a study on the biochronology and biogeography of the Pliocene rhinocerotines from Spain is published by Pandolfi et al. (2022).[127]
Review of the Eocene fossil record of equoids from the Iberian Peninsula is published by Badiola et al. (2022).[128]
New fossil material of palaeotheriids, including the first known records of upper teeth of Franzenium durense and first known mandible and lower teeth of Cantabrotherium, is described from the Eocene (Bartonian) of Mazaterón (Soria, Almazán Basin, Spain) by Perales-Gogenola et al. (2022).[129]
Description of new fossil material of members of the genus Hippotherium from the Miocene of the Linxia Basin (China), providing new information on the skeletal anatomy of members of this genus, and a study on their locomotor capabilities and adaptations to their environment is published by Sun et al. (2022).[130]
A study on the systematic affinities and dietary behavior of Turolian hipparions from the Cioburciu site (Balta Formation; Moldova) is published by Răţoi et al. (2022).[131]
A study on the relationship between size and diet in hipparionins from Vallesian and Turolian circum-Mediterranean localities is published by Orlandi-Oliveras et al. (2022).[132]
Review of the latest occurrences of the hipparions in the Old World, and a study on the taxonomy of the last hipparions is published by van der Made et al. (2022).[133]
Revision of the fossil material of equids from the Khaprovskii Faunal Complex (Russia) is published by Eisenmann (2022).[134]
Revision of the taxonomy of equids from the late Middle Pleistocene to Early Holocene of Apulia (Italy) and a study on their biochronology is published by Mecozzi & Strani (2022).[135]
Revision of the fossil material of Equus stehlini from the Villafranchian of the Upper Valdarno Basin (Tuscany, Italy) is published by Cirilli (2022).[136]

Other laurasiatherians

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Charruatoxodon[137]	Gen. et sp. nov	In press	Ferrero et al.	Late Pliocene–early Pleistocene		Uruguay	A toxodontid notoungulate. The type species is C. uruguayensis.
Diegoaelurus[138]	Gen. et sp. nov	In press	Zack, Poust, & Wagner	Mid Eocene	Santiago Formation	United States ( California)	A machaeroidine oxyaenid. The type species is D. vanvalkenburghae.

Miscellaneous laurasiatherian research

Fragment of a mandible of a notoungulate belonging to the group Interatheriinae is described from the Messinian to Zanclean Tunuyán Formation (Argentina) by Vera & Romano (2022), representing the first record of an interatheriine from this formation and the youngest record of this group reported to date.[139]
A study on the morphological tooth variation in Tremacyllus and on its systematic significance is published by Armella et al. (2022), who recognize Tremacyllus incipiens as a valid taxon.[140]
A study on the postcranial anatomy and likely locomotion of Patriofelis ulta, based on data from two partial skeletons, is published by Kort et al. (2022).[141]
Flink & Werdelin (2022) reconstruct digital endocasts of Quercygale angustidens and Gustafsonia cognita, and evaluate the implications of their anatomy for the knowledge of the evolution of the brain at the origin of Carnivora.[142]

Xenarthrans

Cingulata

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Kelenkura[143]	Gen. et sp. nov	Valid	Barasoain et al.	Late Miocene		Argentina	A glyptodont. The type species is K. castroi.

Cingulatan research

A study investigating the rates of morphological evolution of the skulls of the glyptodonts is published by Machado, Marroig & Hubbe (2022).[144]
Description of new fossil material of Utaetus buccatus from the Eocene Guabirotuba Formation (Brazil), expanding known geographic distribution of this species and representing the first record semi-movable osteoderms in this species reported to date, is published by Sedor et al. (2022).[145]

Pilosa

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images

Pilosan research

A study on the pathological modifications on three articulated vertebrae of a specimen of Eremotherium laurillardi from the Toca das Onças cave (Brazil), and on their implications for the knowledge of the likely cause of death of the animal and on the incorporation mode of skeletal remains into the cave in general, is published by Barbosa et al. (2022).[146]

Other eutherians

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Cokotherium[147]	Gen. et sp. nov		Wang et al.	Early Cretaceous	Jiufotang Formation	China	A basal eutherian. The type species is C. jiufotangensis.
Indoclemensia[148]	Gen. et sp. nov	In press	Mantilla et al.	Paleocene		India	An indeterminate eutherian. Includes the type species I. naskalensis and I. magnus.

Metatherians

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Gumardee keari[149]	Sp. nov	Valid	Travouillon et al.	Miocene		Australia	A member of Macropodiformes.
Gumardee webbi[149]	Sp. nov	Valid	Travouillon et al.	Miocene		Australia	A member of Macropodiformes.

Metatherian research

A study aiming to determine whether it is possible to identify the position of isolated sparassodont teeth using linear discriminant analysis is published by Engelman & Croft (2022).[150]
A study on the evolution and likely causes of extinction of sparassodonts is published by Tarquini, Ladevèze & Prevosti (2022).[151]
A study on the origination and extinction rates of sparassodonts, aiming to determine the cause of their extinction, is published by Pino et al. (2022).[152]
A study on resistances of pedal bones of sthenurine and macropodine kangaroos to bending and cortical bone distribution, and on their implications for the knowledge of possible differences in locomotion of these kangaroos, is published by Wagstaffe et al. (2022).[153]

Prototherians

Name	Novelty	Status	Authors	Age	Type locality	Country	Notes	Images
Murrayglossus[154]	Gen. et comb. nov	Valid	Flannery et al.	Pleistocene		Australia	An echidna. The type species is "Zaglossus" hacketti Glauert (1914).

Other mammals

Name	Novelty	Authors	Age	Country	Notes
Beckumia[155]	Gen. et sp. nov	Martin et al.	Early Cretaceous (Barremian-Aptian)	Germany	A member of Dryolestidae. The type species is B. sinemeckelia.
Cifellitherium[155]	Gen. et sp. nov	Martin et al.	Early Cretaceous (Barremian-Aptian)	Germany	A spalacotheriid symmetrodont. The type species is C. suderlandicum.
Minutolestes[155]	Gen. et sp. nov	Martin et al.	Early Cretaceous (Barremian-Aptian)	Germany	A member of Dryolestidae. The type species is M. submersus.

Other mammalian research

A mammalian petrosal is described from the Lower Cretaceous (Berriasian–Barremian) Batylykh Formation at Teete locality (Sakha, Russia) by Schultz et al. (2022), who tentatively interpret this petrosal as likely to be of eutriconodontan origin.[156]
Second specimen of Corriebaatar marywaltersae, providing new information on the anatomy of this species and confirming its multituberculate affinities, is described from the Early Cretaceous Flat Rocks fossil site (Eumeralla Formation, Australia) by Rich et al. (2022).[157]
Description of new fossil material of Barbatodon oardaensis from Romania is published by Solomon et al. (2022).[158]
Description of a new specimen of Lactodens sheni from the Lower Cretaceous Jiufotang Formation (China), and a study comparing the morphology of the mandible and teeth of this species and Origolestes lii, is published by Mao, Liu & Meng (2022).[159]
A study on the mastication of Peligrotherium tropicalis is published by Harper, Adkins & Rougier (2022).[160]

General research

A study on the phylogenetic relationships of extant and fossil mammals, including previously untested fossils from the Cretaceous-Paleogene transition, is published by Velazco et al. (2022), who recover a new eutherian sister group to Placentalia, and recover Deltatheridium as a marsupial, extending the minimum age of Marsupialia before the Cretaceous-Paleogene boundary.[161]
A study on the evolution of the brain size relative to the body size in placental mammals after the Cretaceous–Paleogene extinction event is published by Bertrand et al. (2022).[162]
A study on the evolution of terrestrial carnivorous mammal diversity in Europe during the Paleogene is published by Solé et al. (2022).[163]
New fossil material of Lagopsis penai and a member of the genus Cainotherium belonging or related to the species C. huerzeleri is described from the Miocene Ribesalbes-Alcora Basin (Spain) by Crespo et al. (2022), who compare the relative abundance of Miocene cainotheriids and lagomorphs in the area, and discuss possible direct interaction between members of both groups.[164]
A study on the environmental variability in Africa during the Pliocene and Pleistocene, and on the impact of this environmental variability on the evolution of African mammals, is published by Cohen et al. (2022).[165]
New marine mammal assemblage, including the youngest pre-Pleistocene earless seal record in South America, is described from the Pliocene Horcón Formation (Chile) by Benites-Palomino et al. (2022).[166]
A study aiming to determine whether the ungulate community associated with Australopithecus afarensis at the Pliocene site of Laetoli (Tanzania) shares similarities with extant communities, and evaluating the implications of this ungulate community for the knowledge of the paleoecology of A. afarensis, is published by Fillion, Harrison & Kwekason (2022).[167]
Systematic description of the Early Pleistocene large mammal fauna from the Maka’amitalu basin (lower Awash Valley, Ethiopia) is published by Rowan et al. (2022).[168]
A study on the equid and suid fossil material from the Early Pleistocene site of Palan-Tyukan (Azerbaijan), and on the implications of these fossils for paleoenvironmental reconstructions, is published by Iltsevich & Sablin (2022).[169]
A study on the foraging ecology of mammals, including early Gigantopithecus blacki, from the Early Pleistocene of the Liucheng Gigantopithecus Cave (Guangxi, China), as indicated by calcium isotope data, is published by Hu et al. (2022).[170]
Revision of the Middle Pleistocene mammalian fauna from the Oumm Qatafa Cave in Palestine, and a study on the implications of this fauna for paleoenvironmental reconstructions, is published by Marom et al. (2022).[171]
A study on the fossil material of reindeers and rodents from the Jankovich Cave and Rejtek I Rock Shelter and on the fossil material of woolly mammoths from the Carpathian Basin (Hungary) is published by Magyari et al. (2022), who evaluate the hypothesis that rapid climate change during the last glacial termination was briefly optimal for grazing megafauna, but these brief optima were followed by rapid regional extinctions, and attempt to determine the order of faunistic and vegetation biome changes in East-Central Europe and its casual linkage.[172]
A study aiming to determine whether brain size was a significant correlate of probability of extinction in Late Quaternary mammals is published by Dembitzer et al. (2022).[173]
A study aiming to determine whether some places, times and types of environment gave rise to abnormal numbers of new species of mammals, based on data from Late Cenozoic fossil record of mammals in Europe, is published by Toivonen, Fortelius & Žliobaitė (2022).[174]
A study on the individual dietary preferences of herbivorous mammals from the Miocene to the present, aiming to determine whether herbivorous generalist species were composed of generalist or specialist individuals, is published by DeSantis et al. (2022).[175]
Gibert et al. (2022) present a spatio-temporal framework that can be used to examine spatial dynamics of Neogene and Pleistocene Old World mammalian communities.[176]

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Wilson Mantilla, Gregory P.; Renne, Paul R.; Samant, Bandana; Mohabey, Dhananjay M.; Dhobale, Anup; Tholt, Andrew J.; Tobin, Thomas S.; Widdowson, Mike; Anantharaman, S.; Dassarma, Dilip Chandra; Wilson Mantilla, Jeffrey A. (2022-02-02). "New mammals from the naskal intertrappean site and the age of India's earliest eutherians". Palaeogeography, Palaeoclimatology, Palaeoecology. 591: 110857. Bibcode:2022PPP...591k0857W. doi:10.1016/j.palaeo.2022.110857. ISSN 0031-0182. S2CID 246508839.
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